This was the bit that caught my eye the most:
‘Large-scale culls of mountain hares to reduce tick loads, in order to benefit grouse and other bird survival, will only be effective when other hosts are absent, or their ability to host ticks are similarly reduced. This will not be the case for many estates in Scotland.
On the basis of the available evidence, there is no compelling field evidence for undertaking large-scale mountain hare culls to control Louping Ill Virus in areas of Scotland where there are high densities of other tick-bearing mammals. Culls should therefore not be undertaken for this purpose in these circumstances.‘